Season 1 (Classic): The biggest twist is that the goofy protagonist Usagi is the Moon Princess everyone is searching for. The finale kills off all main characters, only to resurrect them with amnesia—a shocking move for a kids' show.
Season 2 (R): Introduces Chibiusa, who is revealed to be Usagi and Mamoru's future daughter from the 30th Century, turning the romance into a timeline-spanning dynasty.
Season 3 (S): Reveals that the 'Messiah of Silence' destined to destroy the world is the frail girl Hotaru Tomoe (Sailor Saturn). The twist is that Usagi saves the world not by killing her, but by allowing her to awaken and reset the world peacefully.
Season 5 (Stars): The ultimate villain, Sailor Galaxia, is revealed to be a fallen hero—the strongest Sailor Guardian who sealed Chaos within herself to save the universe but was eventually corrupted by it. The series ends with Usagi embracing Chaos itself, accepting that light and dark must coexist.