Following his transition from world-conquering supervillain to devoted adoptive father, Felonious Gru has settled into an idyllic domestic life. Alongside his loyal cadre of Minions and his aging cohort Dr. Nefario, Gru focuses on domestic duties, organizing elaborate birthday celebrations for Margo, Edith, and Agnes while attempting to pivot their underground laboratory into an artisanal jam and jelly manufacturing business.
Gru's peaceful suburban existence is disrupted when he is abruptly abducted by Agent Lucy Wilde and brought to the submerged headquarters of the Anti-Villain League (AVL). AVL Director Silas Ramsbottom requests Gru's unique criminal perspective to track down an elusive mastermind who stole the volatile PX-41 mutagen from a secret Arctic laboratory. Despite his initial reluctance and Dr. Nefario's departure to seek more malevolent employment, Gru agrees to go undercover at the Paradise Shopping Mall posing as a bakery proprietor alongside the quirky, enthusiastic Lucy.
While staking out suspicious shop owners, including flamboyant salsa restaurateur Eduardo Pérez, Gru faces unexpected complications. He must juggle his protective fatherhood over Margo's first adolescent crush, his emerging romantic feelings for Lucy, and the mysterious, sudden disappearances of his beloved Minions, forcing him to decide what kind of hero he truly wants to become.
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