In a world strictly divided by fear and prejudice, bears live in the city above ground while mice dwell in the sewers below. Celestine, an orphan mouse with an artistic soul, is forced to collect bear teeth for her society's dental industry but dreams of drawing. She constantly questions the terrifying tales told about bears. Meanwhile, Ernest, a destitute and solitary bear living on the fringes of society, wakes up from hibernation starving and grumpy.
Their paths collide when Ernest finds Celestine in a trash can. Instead of eating her, as social norms dictate, he strikes a bargain: she helps him break into a candy shop, and he helps her steal a sack of teeth. This crime spree turns them into fugitives hunted by the police of both worlds. Taking refuge in Ernest's secluded cabin during the winter, they overcome their mutual prejudices and bond over their shared love for music and art, forming an unlikely family.
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